Gouania adriatica

Wagner, Kovačić and Koblmüller 2020



Gouania wildenowi  in part

Previously referred to by the informal name of Adriatic stout Gouania (Wagner et al. 2019).




Holotype: PMR VP4618, male, 41.41+6.77 mm

Paratypes: ZSM-PIS-047650, male, 41.03+5.89 mm; ZSM-PIS-047652, male, 27.3+4.27 mm; ZSM-PIS-047651, male, 34.53+5.65 mm ;PMR VP3523, juvenile of unidentified sex, 22.95+3.6 mm; PMR VP3524, male. 26.19 mm, caudal fin damaged and PMR VP3525, female, 28.4+4.23 mm; ZSM-PIS-047653, female, 34.42+4.85 mm and ZSM-PIS-047653, female, 35.23+4.9 mm; PMR VP4580, female, 34.7+4.99 mm


Type locality: Stoja, Pula, Croatia, 44°51'38.4"N, 13°49'05.0"E.

Known also from:

Envi beach, Vlorë, Albania, 40°23'16.3"N 19°28'58.2"E

Trstenik Pelješac, Croatia, 42°54'07.7"N 17°25'47.0"E

Stara Baška, Krk, Croatia, 44°56'45.3"N 14°42'22.2"E

Glavotok, Krk, Croatia, 45°05'44.9"N 14°26'32.4"E

Pećine, Rijeka, Croatia, 45°18'52.4"N 14°28'11.7"E

Sv. Marina, Istria, Croatia, 45°01'42.1"N 14°09'17.4"E


A member of the Subfamily Lepadogastrinae together with the following genera:  Apletodon (7 species), Diplecogaster (8 species), Lepadogaster (3 species), Lecanogaster (2 species) and Opeatogenys (2 species) (Briggs 1957).

Conservation Status


Internet Resources

Key References

Risso, A. Book 1810 Ichthyologie de Nice, ou histoire naturelle des poissons du Département des Alpes Maritimes
Nardo, G.D. Journal Article 1827 Prodromus observationum et disquisitionum Adriaticae ichthyologiae
Nardo, G.D. Journal Article 1827 Prodromus observationum et disquisitionum ichthyologiae Adriaticae
Nardo, G.D. Journal Article 1833 Eine neue Art von Lepadogaster (L. piger)
Swainson, W. Book 1839 On the natural history and classification of fishes, amphibians, & reptiles, or monocardian animals. Volume 2
Troschel, F.H. Journal Article 1860 Leptopterygius, neue Gattung der Discoboli
Briggs, J.C. Journal Article 1955 A monograph of the clingfishes (order Xenopterygii)
Leray, C. Journal Article 1961 Contribution a l'étude ostéologique de Gouania wildenowii Risso (Téléostéens) (squelette céphalique et ceintures)
Peres, J.M. Journal Article 1967 The Mediterranean benthos
Katayama, M and Ikeda, H Journal Article 1972 Luciogobius pallidus collected at Nagato-shi in Yamaguchi Prefecture
Birdsong, R.S., Murdy, E.O. and Pezold, F. Journal Article 1988 A study of the vertebral column and median fin osteology in gobioid fishes with comments on gobioid relationships.
Hofrichter, R. Thesis 1995 Taxonomie, Verbreitung und Ökologie von Schildfischen der Unterfamilie Lepadogastrinae (Gobiesocidae, Teleostei)
Herler, J.R.A., Patzner, A.A. and Hilgers, H. Journal Article 1999 Habitat selection and ecology of two speleophilic gobiid fishes (Pisces: Gobiidae) from the western Mediterranean Sea
Patzner, R. Journal Article 1999 Habitat utilization and depth distribution of small cryptobenthic fishes (Blenniidae, Gobiesocidae, Gobiidae, Tripterygiidae) in Ibiza (western Mediterranean Sea).
Hofrichter, R. and Patzner, R.A. Journal Article 2000 Habitat and microhabitat of Mediterranean clingfishes (Teleostei: Gobiesociformes: Gobiesocidae)
Almada, F., Henriques, M., Levy, A., Pereira, A.M., Robalo, J.I. and Almada, V.C. Journal Article 2008 Reclassification of Lepadogaster candollei based on molecular and meristic evidence with a redefinition of the genus Lepadogaster
Maeda, K., Yamasaki, N., Kondo, M. and Tachihara, K. Journal Article 2008 Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island
Yamada, T., Sugiyama, K, Tamaki, K, Kawakita, A. and Kato, M. Journal Article 2009 Adaptive radiation of gobies in the interstitial habitats of gravel beaches accompanied by body elongation and excessive vertebral segmentation
Ellingson, R.A. Thesis 2012 Phylogenetics and phylogeography of North Pacific bay gobies: adaptive convergence, relictual endemism, and climate-driven population structure
Ellingson, R.A., Swift, C.C., Findley, L.T. and Jacobs, D.K. Journal Article 2014 Convergent evolution of ecomorphological adaptations in geographically isolated Bay gobies (Teleostei: Gobionellidae) of the temperate North Pacific
Bilecenoglu, M. Journal Article 2015 First observation of the amphibious behaviour of Gouania willdenowi (Gobiesocidae) in the Eastern Mediterranean Sea
Wagner, M., Bračun, S. and Koblmüller, S. Conference Paper 2017 Cryptic diversity in pebble beaches: Rapid speciation and convergent evolution in the genus Gouania (Gobiesocidae)
King, C.D. Thesis 2017 Molecular phylogenetic investigation of the clingfishes (Teleostei: Gobiesocidae)
Wagner, M. Thesis 2018 New insights into the phylogeography and evolutionary history of two Mediterranean clingfish genera (Teleostei, Gobiesocidae)
Wagner, M., Bračun, S., Skofitsch, G., Kovačić, M., Zogaris, S., Iglésias, S.P., Sefc, K.M. and Koblmüller, S. Journal Article 2019 Diversification in gravel beaches: A radiation of interstitial clingfish (Gouania, Gobiesocidae) in the Mediterranean Sea
Kondo, K. and Kato, M. Journal Article 2019 Life in coastal pebble sediment: Unique interstitial organism community and selective feeding on meiobenthos by interstitial fishes (Luciogobius: Gobiidae)
Mirete-Bachiller, S., Olivieri, D.N., Gambón-Deza, F. Journal Article 2019 Gouania willdenowi is a teleost fish without immunoglobulin genes
Yun, S.-M., Park, J.-M. and Han, K.-H. Journal Article 2020 Osteological development of the larvae and juvenile of Luciogobius grandis (Pisces: Gobiidae)
Charmpila, E.A., Wagner, M. and Reichenbacher, B. Journal Article 2021 First documentation of the otoliths of the species of Gouania (Teleostei: Gobiesocidae) in the Mediterranean Sea.
Wagner, M., Kovacic, M. and Koblmüller, S. Journal Article 2021 Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra.