Subterranean Fishes of the World

Coexisting Species

Discussions of the coexistence of cave fish species can be found in Anonymous (1976), Smith (1978), Thinès (1978) and Borowsky and Mertz (2001).

Notes on the coexistence of subterranean fishes

The total number of species in pairs and triplets is 58, which is a remarkable ~23% of all known species (247 in March 2019).

^Tb indicates a troglobiotic species, ^EuT a eutroglophile species.

Location	Species 1	Species 2/3	References
USA (Kentucky)	Amblyopsis spelaea ^Tb	Typhlichthys subterraneus ^Tb	Eigenmann, C.H. (1909) Woods, L.P. and Inger, R.F. (1957) Barr, T.C. (1967) Barr, T.C. and Kuehne, R.A. (1971)
USA (Texas)	Trogloglanis pattersoni ^Tb	Satan eurystomus ^Tb	Suttkus, R. D. (1961) Mitchell, R.W. (1969) Longley, G. (1979)
Mexico (Yucatan)	Typhlias pearsei ^Tb	Ophisternon infernale ^Tb	Reddell, J. R. (1981) Wilkens, H. (1982)
Mexico (Acàtlan)	Rhamdia reddelli ^Tb	Rhamdia zongolicensis ^Tb	Mosier, D. (1984)
Cuba (Habana)	Lucifuga dentata ^Tb	Lucifuga subterranea ^Tb	Eigenmann, C.H. (1909) Nalbant, T. T. (1981) Diaz Perez, P.A. (1988)
Cuba (Matanzas)	Lucifuga simile ^Tb	Lucifuga dentata ^Tb	Burgess, G.H. and Franz, R. (1987)
India (Kerala)	Horaglanis krishnai ^Tb	Rakthamichthys indicus ^Tb	Eapen, K. C. (1963)
Iraq (Haditha)	Garra widdowsoni ^Tb	Caecocypris basimi ^Tb	Banister, K.E. and Bunni, M.K. (1980)
Iran (Kaaje Ru)	Garra typhlops ^Tb	Eidinemacheilus smithi ^Tb Garra lorestanensis ^Tb	Greenwood, P. H. (1976) Anonymous (1976) Thines, G. (1978) Smith, A. (1978) Mousavi-Sabet, H. and Eagderi, S. (2016) Vatandoust, S., Mousavi‐Sabet, H., Geiger, M.F. and Freyhof, J. (2019)
Australia (NW Cape)	Milyeringa veritas ^Tb	Ophisternon candidum ^Tb	Mees, G.F. (1962) Humphreys, W. F. and Adams, M. (1991)
Thailand	Nemacheilus troglocataractus ^Tb	Pterocryptis buccata ^Tb	Gery, J. (1987) Kottelat, M. (1998) Ng, H.H. and Kottelat, M. (1998)
Thailand	Cryptotora thamicola ^Tb	Schistura oedipus ^Tb	Borowsky, R. (1998)
Thailand (Phitsanulok)	Neolissochilus subterraneus ^Tb	Schistura spiesi ^Tb	Smart, D. (1999) Borowsky, R.L. and Vidthayanon, C. (2001) Vidthayanon, C. and Kottelat, M. (2003)
Brazil (Goias)	Ancistrus cryptophthalmus ^Tb	Ituglanis passensis ^Tb	Trajano, E. and de Souza, A. M. (1994)
China (Yunnan)	Sinocyclocheilus anophthalmus ^Tb	Triplophysa yunnanensis	Chen, Y.R. and Yang, J.X. (1993)
China	Sinocyclocheilus furcodorsalis ^Tb	Triplophysa tianeensis ^Tb	Romero, A., Zhao, Y.H. and Chen, X.Y. (2009)
China	Sinocyclocheilus lingyunensis ^Tb	Troglonectes lingyunensis ^Tb Sinocyclocheilus microphthalmus ^Tb	Romero, A., Zhao, Y.H. and Chen, X.Y. (2009) Li, WX, Xiao, H, Zan, RG, Luo, ZF and Li, HG (2000)
China	Sinocyclocheilus lingyunensis ^Tb	Sinocyclocheilus microphthalmus ^Tb Troglonectes lingyunensis ^Tb
China	Sinocyclocheilus longifinus ^Tb
China	Triplophysa macrocephala ^Tb	Troglonectes longibarbatus ^Tb
China	Bibarba parvoculus ^Tb	Karstsinnectes anophthalmus Sinocyclocheilus brevis ^Tb	Wu, T.J., Yang, J. and Xiu, L.H. (2015)
USA (Alabama)	Typhlichthys undescribed species lineage Key Cave ^Tb	Speoplatyrhinus poulsoni ^Tb	U.S. Fish and Wildlife Service (1991)
China	Sinocyclocheilus hyalinus ^Tb	Triplophysa aluensis
China	Protocobitis typhlops ^Tb	Troglonectes translucens ^Tb	Chen, Y.R. and Yang, J.X. (1993) Yang, J.X., Chen, Y.R. and Lan, J.H. (1994)
China	Sinocyclocheilus tianlinensis ^Tb	Triplophysa tianlinensis ^Tb	Li, J., Li, X.H., Lan, J.H. and Du, L.N. (2016)
Madagascar	Typhleotris madagascariensis ^Tb	Typhleotris mararybe ^Tb	Sparks, J.S. and Chakrabarty, P. (2012)
Madagascar	Typhleotris pauliani ^Tb	Typhleotris mararybe ^Tb	Sparks, J.S. and Chakrabarty, P. (2012)
Australia (Barrow island)	Milyeringa justitia ^Tb	Ophisternon candidum ^Tb	Larson, H.K., Foster, R., Humphreys, W.F. and Stevens, M. (2013)
Brazil	Copionodon undescribed species Igatu ^Tb	Glaphyropoma spinosum ^Tb	Bichuette, M.E., de Pinna, M.C.C. and Trajano, E. (2008)
China	Triplophysa rosa ^Tb	Triplophysa wulongensis ^Tb	Chen, X.Y. and Yang, J. (2005) Chen, S., Sheraliev, B., Shu, L. and Peng, Z. (2021)
Philippines	Barbodes pyrpholeos	Caecogobius personatus	Larson, H.K. and Husana, D.E.M. (2018) Husana, D.E.M. (2020) Tan, H.H. and Husana, D.E.M. (2021)